definition
An undifferentiated plant body, such as in algae.
definition
Any plant body lacking vascular tissue.
definition
An undifferentiated plant body, such as in algae.
definition
Any plant body lacking vascular tissue.
The Mosses and Liverworts include forms with a more or less leaf-like thallus, such as many of the liverworts, and forms in which the plant shows a differentiation into a stem bearing remarkably simple leaves, as in the true mosses.
This process may result in a considerable thickening of the thallus.
Thallophyta are the most lowly organized plants and include a great variety of forms, the vegetative portion of which consists of a single cell or a number of cells forming a more or less branched thallus.
The thallus in all cases consists of a branched filament of cells placed end to end, as in many of the Green Algae.
Or the thallus may have a leaf-like form, the branches from the central threads which form the midrib growing out mainly in one plane and forming a lamina, extended right and left of the midrib.
In many Laminariaceae the thallus also grows regularly in thickness by division of its surface layer, adding to the subjacent permanent tissue and thus forming a secondary meristem.
In the Bryophytes water is still absorbed, not only from the soil but also largely from rain, dew, &c., through the general surface of the subaerial body (thallus), or in the more differentiated forms through the leaves.
A sufficient description of the thallus of the liverworts will be found in the article BRYOIHYTA.
The frondose (thalloid) Jungermanniales show no such differentiation of an assimilating tissue, though the upper cells of the thallus usually have more chlorophyll than the rest.
They serve to conduct water through the thallus, the assimilating parts of which are in these forms often raised above the soil and are comparatively remote from the rhizoid-bearing (water-absorbing) region.
In the liverworts we find fixation of the thallus by water-absorbing rhizoids; in certain forms with a localized region of water-absorption the development of a primitive hydrom or water-conducting system; and in others with rather a massive type of thallus the differentiation of a special assimilative and transpiring system.
The gametophyte, which bears the sexual organs, is either a free-living thallus corresponding in degree of differentiation with the lower liverworts, or it is a mass of cells which always remains enclosed in a spore and is parasitic upon the sporophyte.
From the primitive uniform Systems. mass of undifferentiated assimilating cells, which we may conceive of as the starting-point of differentiation, though such an undifferentiated body is only actually realized in the thallus of the lower Algae, there is, (1) on the one hand, a specialization of a surface layer regulating the immediate relations of the plant with its surroundings.
In mounting collemas it is advisable to let the specimen become dry and hard, and then to separate a portion from adherent mosses, earth, &c., and mount it separately so as to show the branching of the thallus.
The green (or blue-green) cells were termed gonidia by Wallroth, who looked upon them as asexual reproductive cells, but when it was later realized that they were not reproductive elements they were considered as mere outgrowths of the hyphae of the thallus which had developed chlorophyll.
Rees in 1871 produced the sterile thallus of a Collema from its constituents; later Stahl did the same for three species.
The thallus or body of the lichen is of very different form in different genera.
We can distinguish this class of forms as lichens with a homoiomerous thallus, i.e.
The majority of the lichens, however, possess a stratified thallus in which the gonidia are found as a definite layer or layers embedded in a pseudoparenchymatous mass of fungal hyphae, i.e.
In external form the heteromerous thallus presents the following modifications.
I, Germinating ascospore (sp) 2, Thallus in process of formawith branching germ-tube tion.
In colour also the thallus externally is very variable.
The surface of the thallus often exhibits outgrowths in the form of warts, hairs, &c. The medullary layer, which usually forms the main part of the thallus, is distinguished from the cortical layer by its looser consistence and the presence in it of numerous, large, air-containing spaces.
These structures are known as cephalodia and they usually occupy a definite position in the thallus.
They are distinguished by possessing as gonidia algae foreign to the ordinary part of the thallus.
In the first process, portions of thallus containing gonidia may be accidentally separated and so may start new plants.
Since they are provided with both fungal and algal elements, they are able to develop directly, under suitable conditions, into a new thallus.
The soredia are the most successful method of reproduction in lichens, for not only are some forms nearly always without spore-formation and in others the spores laregly abortive, but in all cases the spore represents only the fungal component of the thallus, and its success in the development of a new lichen-thallus depends on the chance meeting, at the time of germination, with the appropriate algal component.
In the other view the spermatia are the male sexual cells and thus A, Optical longitudinal section of the ex are rightly named; it tremity of a thin branch of the thallus should, however, be which has become transparent in pointed out that this solution of potash.
In favour of the conidial view is the fact that in the case of Collema and a few other forms the spermatia have been made to germinate in artificial cultures, and in the case of Calicium parietinum Moller succeeded in producing a spermogonia bearing thallus from a spermatium.
For the germination of the spermatia in nature there is only the observation of Hedlund, that in Catillaria denigrata and C. prasena a thallus may be derived from the spermatia under natural conditions.
The apothecia, though of the normal fungal type and usually disk-shaped, are somewhat more variable, and since the Morphologie and Biologie der Pilze, Mycetozoen und 1 The thalline margin (margo thallinus) is the projecting edge of a special layer of thallus, the amphithecium, round the actual apothecium; the proper margin (margo proprius) is the projecting edge of the apothecium itself.
He showed that the archicarp consisted of two parts, a lower coiled portion, the ascogonium, and an upper portion, the trichogyne, which projected from the thallus.
When the spores germinate the germ-tubes surround the algal cells, which now increase in size and become the normal gonidia of the thallus.
Clavaria mucida, however, h a s apparently some claims to be considered as a Basidiolichen, since the base of the fruit body and the thallus from which it arises, according to Coker, always shows a mixture of hyphae and algae.
When the fungus predominates in the thallus it has a bracket-like mode of growth and is found projecting from the branches of trees with the hymenium on the under side.
A solution of iodine is also used as a test owing to the blue or wine-red colour which the thallus, hymenium or spores may give with this reagent.
Most systems agree in deriving the major divisions from the characters of the reproductive organs (perithecia, apothecia, or basidiospore bearing fructification), while the characters of the algal cells and those of the thallus generally are used for the minor divisions.
Fungi with segmental thallus; sexual reproduction sometimes with typical antheridia and oogonia (ascogonia) but usually much reduced.
Forms with septate thallus, and reproduction by chlamydospores which on germination produce sporidia; sexuality doubtful.
Thallus septate; spores developed in special type of sporangium, the ascus, the number of spores being usually eight.
Amongst the lower plants, however, it is found that a demarcation into stem and leaf is impossible, but that there is a structure which partakes of the characters of both - such is a thallus.
Certain species are regularly found in the intercellular spaces of higher plants; such are species of Nostoc in the thallus of Anthoceros, the leaves of Azolla and the roots of Cycads.
The thallus may be unicellular or multicellular.
Valoniaceae and The thallus is of more varied structure in this group than in any Dasycladaceae are partially septate, but elsewhere no cellulose partiother.
In the simplest case it may consist of a single cell, which may tions occur, and the thallus is more or less the continuous tube from remain free during the whole of the greater part of its t xistence, which the group is named.
The Ulvaceae, the thallus of which consists of external form as an expanded Coprinus, Neomeris simulates the laminae, one or more cells thick, or hollow tubes, probably represent fertile shoot of Equisetum with its densely packed whorled branches, a still more advanced stage in the passage of a colony into a multiand in Microdictyon, Anadyomene, Struvea and Boodlea the branches, cellular plant.
Attached to the bottom of pools series of the Confervales, the thallus consists of filaments branched by means of rhizoids, the thallus of Characeae grows upwards by or unbranched, attached at one extremity, and growing almost means of an apical cell, giving off whorled appendages at regular wholly at the free end.
There is thus a Protococcales, and in the bulk of the Confervales, the thallus consists close approach to the external morphology of the higher plants.
Valonia and Caulerpa among Siphonales detach portions of their thallus, which are capable of independent growth.
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